iPad Screenshots


TimeTree 4 is a public knowledge-base for information on the evolutionary timescale of life. This application allows easy exploration of the thousands of divergence times among organisms in the scientific literature. A tree-based (hierarchical) system is used to identify all published molecular time estimates bearing on the divergence of two chosen organisms, such as species, compute summary statistics, and present the results. Names of two taxa to be compared are entered in the search window and the results are presented on a set of self-explanatory tabs.

TimeTree 4 contains data from 2,274 studies including 50,632 species. We will be adding more data in the future as it comes in from researchers.

TimeTree 4 requires iOS 8.0 or newer.

The TimeTree project is jointly directed by Blair Hedges (Center for Biodiversity, Temple University) and Sudhir Kumar (Institute for Genomics and Evolutionary Medicine, Temple University). This project has been supported, in part, by grants from the National Science Foundation, National Institutes of Health, NASA Astrobiology Institute, and Science Foundation of Arizona.

What’s New

Version 4.0

New universal app for iPad and iPhone.
New divergence times from over 2000 studies.
Numerous bug fixes and server updates.

Ratings and Reviews

4.6 out of 5
5 Ratings

5 Ratings

d1s80 ,


wonderful app that reaches further and further with citations and data. Comes from a world renown evolution genetics program. All I can say is thanks

HJKarten ,

Please correct my previous review

I just submitted a generally favorable review of this APP, but complained that it needed a version specifically adapted to the iPad/iPad2. I then went back to the APP store and realized that there are two separate versions, one for the iTouch/iPhone, and a second version for the iPad. The iPad version uses full screen, but only displays in portrait mode, not landscape.
Very useful program.

Nariokotome ,


Functions without issue. Data are accurate with references linked to NCBI summaries. Great reference and useful for illustrating evolutionary relationships in class. Bravo!


Center for Evolutionary Functional Genomics
68.6 MB

Requires iOS 8.0 or later. Compatible with iPad.



Age Rating


  • Family Sharing

    With Family Sharing set up, up to six family members can use this app.

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