iPad Screenshots


TimeTree 4 is a public knowledge-base for information on the evolutionary timescale of life. This application allows easy exploration of the thousands of divergence times among organisms in the scientific literature. A tree-based (hierarchical) system is used to identify all published molecular time estimates bearing on the divergence of two chosen organisms, such as species, compute summary statistics, and present the results. Names of two taxa to be compared are entered in the search window and the results are presented on a set of self-explanatory tabs.

TimeTree 4 contains data from 2,274 studies including 50,632 species. We will be adding more data in the future as it comes in from researchers.

TimeTree 4 requires iOS 8.0 or newer.

The TimeTree project is jointly directed by Blair Hedges (Center for Biodiversity, Temple University) and Sudhir Kumar (Institute for Genomics and Evolutionary Medicine, Temple University). This project has been supported, in part, by grants from the National Science Foundation, National Institutes of Health, NASA Astrobiology Institute, and Science Foundation of Arizona.

What’s New

Version 4.0

This app has been updated by Apple to use the latest Apple signing certificate.

New universal app for iPad and iPhone.
New divergence times from over 2000 studies.
Numerous bug fixes and server updates.

Ratings and Reviews

1.0 out of 5
1 Rating

1 Rating

HJKarten ,

Please correct my previous review

I just submitted a generally favorable review of this APP, but complained that it needed a version specifically adapted to the iPad/iPad2. I then went back to the APP store and realized that there are two separate versions, one for the iTouch/iPhone, and a second version for the iPad. The iPad version uses full screen, but only displays in portrait mode, not landscape.
Very useful program.

d1s80 ,


wonderful app that reaches further and further with citations and data. Comes from a world renown evolution genetics program. All I can say is thanks

Nariokotome ,


Functions without issue. Data are accurate with references linked to NCBI summaries. Great reference and useful for illustrating evolutionary relationships in class. Bravo!

App Privacy

The developer, Arizona and Penn State Universities, has not provided details about its privacy practices and handling of data to Apple.

No Details Provided

The developer will be required to provide privacy details when they submit their next app update.


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